"Our lust is brief. We are means to those small creatures within us, and nature has other ends than we."
I believe that Joyce was referring to the brevity of life and the whole of nature and the existence of a consciousness related to that whole. Humans are only a part of the whole of the fauna and flora. We're not the purpose of nature. Nature is not an entity that exists outside of us for our exploitation. We are a small, but exceptional part of it. But, that is all. In the end, we rarely get to see what got us. These creatures are just too small.